Causes? I have no idea, but by the fact that he didn't seem to be able to get up, I'm going to assume that he had some kind of a swim bladder issue. If that's even possible.
Every thing else in the tank seems unaffected.
As I'm typing this I notice that I use the word "seems" a lot. I guess I do that because it is just my observation, the way I see it. I'm not sure if that is the fact or not. In this hobby there are so many variables that we do not control. So, unless you have a Lab and a controlled environment where everything is monitored and you do biopsies when something perishes, we are just guessing, even if it is an educated guess.
Here is one of my favorite pictures of him hanging out in the toadstool coral.

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